It is also easy to see that our previous equation can be written as δLj=∂C∂zLj=∂C∂aLj∂aLj∂zLj, which can also be interpreted as the partial derivative of the cost function with respect to the biases bLj, namely δLj=∂C∂bLj∂bLj∂zLj=∂C∂bLj, That is, the error δLj is exactly equal to the rate of change of the cost function as a function of the bias.